Sasaki H, Khalife A, Ito F, 2025. Colony composition, prey preference and behavioral characteristics of Leptanilla kubotai (Hymenoptera: Formicidae: Leptanillinae). Asian Myrmecology 18: 018001 (1-9). https://doi.org/10.20362/am.018001
Khalife A, Sasaki H, Ito F, 2024. Intercastes in the ant Aphaenogaster rugulosa (Formicidae: Myrmicinae): morphological variation and adaptive potential. Insectes Sociaux 71, 377–387. https://doi.org/10.1007/s00040-024-00987-y
Khalife A, Billen J, Economo EP, 2023. Evidence of a thoracic crop in workers, soldiers, and queens of Carebara perpusilla ants (Formicidae: Myrmicinae). The Science of Nature 110, 36. https://doi.org/10.1007/s00114-023-01866-5
Khalife A, Peeters C, Economo EP, 2022. Minute workers and large soldiers in the subterranean ant Carebara perpusilla: musculoskeletal consequences of Haller’s rule in the thorax. Arthropod Structure & Development 69, 101188. https://doi.org/10.1016/j.asd.2022.101188
Billen J, Khalife A, Ito F, Anh ND, Esteves FA, 2021. The basitarsal sulcus gland, a novel exocrine structure in ants. Arthropod Structure & Development 61, 101041. https://doi.org/10.1016/j.asd.2021.101041
Khalife A, Peeters C, 2020. Food storage and morphological divergence between worker and soldier castes in a subterranean myrmicine ant, Carebara perpusilla. Journal of Natural History 54(47-48), 3131-3148. https://doi.org/10.1080/00222933.2021.1890851
Peeters C, Keller RA, Khalife A, Fischer G, Katzke J, Blanke A, Economo EP, 2020. The loss of flight in ant workers enabled an evolutionary redesign of the thorax for ground labour. Frontiers in Zoology 17, 1-13. https://doi.org/10.1186/s12983-020-00375-9
Khalife A, Keller RA, Billen J, Hita Garcia F, Economo EP, Peeters C, 2018. Skeletomuscular adaptations of head and legs of Melissotarsus ants for tunnelling through living wood. Frontiers in Zoology 15, 1-11. https://doi.org/10.1186/s12983-018-0277-6
Lecchini D, Bertucci F, Gache C, Khalife A, Besson M, Roux N, … Hédouin L, 2018. Boat noise prevents soundscape-based habitat selection by coral planulae. Scientific Reports 8(1), 9283. https://doi.org/10.1038/s41598-018-27674-w